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Spergularia marina

(L.) Besser (1822)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-CoreEudicot Order: Caryophyllales Family: Caryophyllaceae Genus: Spergularia

mimi‘ïlio [mimiilio], saltmarsh sand spurry

Description

Key Characters:

Growth Form: Annual herbs, often fleshy and growing in saline areas.

Stems: Stems erect to nearly prostrate, somewhat fleshy, 0.5–3 dm long, unbranched or many-branched, glabrous or glandular puberulent.

Roots:

Leaves: Leaves simple. Opposite or rarely fascicled. Blades linear, 5–40 mm long, 0.6–1.5 mm wide. Apex blunt to apiculate. Base leaf bases often connate around the stem. Surfaces glabrous or glandular puberulent, blades often fleshy. Margins entire. 1-veined. Sessile. Stipules deltate, 2–4 mm long, dry and membranous.

Flowers: Flowers in lax, elongate, racemose cymes. Flowers bisexual (perfect), actinomorphic. Sepals ovate, 2.5–5 mm long; distinct, nearly distinct, or connate into a conspicuous tube. Corolla of 5 petals, petals white to pink, ovate, 2–4 mm long. Stamens 2–5, distinct or adnate to petals at base to form a short tube, which may be adnate to the gynophore or sometimes adnate to lower part of calyx, petaloid staminodes sometimes present; anthers dithecal, opening by longitudinal slits. Ovary superior; styles 3, distinct or nearly so, filiform, 0.2–3 mm, glabrous proximally; stigmas 3, linear along adaxial surface of styles.

Fruit: Capsules 3.5–6 mm long; dehiscing by 3(5) valves. Seeds brown; 0.6–0.8 mm long; smooth or papillose; usually not winged; perisperm starchy; hard or rarely soft; true endosperm absent.

Ploidy: 2n = 18; 34;36

Habitat: Naturalized in coastal and low elevation.

Elevation Range:

Historical Distribution

Synonyms (125)

Alsine atheniensis proles marina (L.) Samp. (1913)
Alsine atheniensis proles urbica (Leffler) Samp. (1913)
Alsine marginata C.A.Mey. (1831)
Alsine marina (L.) Mert. & W.D.J.Koch (1826)
Alsine marina var. succosior Mert. & W.D.J.Koch (1831)
Alsine maritima proles urbica (Leffler) Samp. (1914)
Alsine salina J.Groves (1904)
Alsine sparsiflora (Greene) House (1921)
Alsine tenuis (Greene) House (1921)
Alsinella media Hornem. (1827)
Arenaria halophila Bunge (1830)
Arenaria heterosperma Guss. (1843)
Arenaria marina (L.) Weber (1787)
Arenaria maritima (L.) Steud. (1840)
Arenaria media DC. (1824)
Arenaria miquelonensis Bach.Pyl. ex Lebel (1868)
Arenaria rubra subsp. marina (L.) Ehrh. (1780)
Arenaria rubra var. marina L. (1753)
Arenaria salina (J.Presl & C.Presl) Ser. (1824)
Buda marina (L.) Dumort. (1827)
Buda marina var. minor S.Watson (1890)
Buda media var. marina (Griseb.) Kuntze (1891)
Corion halophilum Merino (1905)
Corion halophilum var. dillenii (Lebel) Merino (1905)
Corion halophilum var. urbicum (Leffler) Merino (1905)
Corion marinum (L.) N.E.Br. (1891)
Corion marinum var. leiospermum (Kindb.) N.E.Br. (1891)
Holosteum sperguloides Lehm. (1828)
Lepigonum caninum Leffler (1869)
Lepigonum caninum var. urbicum Leffler (1869)
Lepigonum leiospermum Kindb. (1863)
Lepigonum marinum (L.) Wahlenb. (1820)
Lepigonum medium Fr. (1842)
Lepigonum neglectum Kindb. (1856)
Lepigonum salinum (J.Presl & C.Presl) G.Don (1839)
Lepigonum salinum var. sperguloides (Lehm.) Leffler (1865)
Lepigonum tenue Greene (1887)
Lepigonum trachyspermum subsp. murale Kindb. (1863)
Spergula canina subsp. sperguloides (Lehm.) Ahlfv. (1901)
Spergula coreana H.Lév. (1913)
Spergula heterosperma (Guss.) E.A.Durand & Barratte (1910)
Spergula marina (L.) Bartl. & H.L.Wendl. (1825)
Spergula salina (J.Presl & C.Presl) D.Dietr. (1840)
Spergula salina subvar. halophila (Bunge) Maire & Weiller (1963)
Spergula salina var. australis (Lebel) Maire (1932)
Spergula salina var. faurei Maire (1940)
Spergula salina var. halophila (Bunge) Maire (1935)
Spergula salina var. leiosperma (Kindb.) Johanss. (1910)
Spergula salina var. longicaulis (Pomel) Maire & Weiller (1963)
Spergula salina var. sperguloides (Lehm.) Maire (1941)
Spergula salina var. stenopetala (A.Chev.) Maire & Weiller (1963)
Spergula salina var. urbicum (Leffler) Maire (1932)
Spergula seminulifera (Hy) Maire (1932)
Spergularia alata Wiegand (1920)
Spergularia bofillii Sennen (1927)
Spergularia bungeana Tzvelev (2000)
Spergularia caballeroi Sennen & Mauricio (1933)
Spergularia campestris var. pinguis (Fenzl) P.Fourn. (1936)
Spergularia dillenii Lebel (1869)
Spergularia dillenii proles salina (J.Presl & C.Presl) Rouy & Foucaud (1896)
Spergularia dillenii subsp. urbica (Leffler) Rouy & Foucaud (1896)
Spergularia dillenii var. australis Lebel (1869)
Spergularia dillenii var. confusa Rouy & Foucaud (1896)
Spergularia dillenii var. pedicellaris Rouy & Foucaud (1896)
Spergularia dillenii var. perennis Rouy & Foucaud (1896)
Spergularia fauriei (Nakai) Nakai (1952)
Spergularia halophila T.Marsson (1869)
Spergularia halophila var. dillenii (Lebel) Samp. (1904)
Spergularia halophila var. marina (L.) T.Marsson (1869)
Spergularia halophila var. urbica (Leffler) Samp. (1904)
Spergularia heterosperma Lebel (1869)
Spergularia leiosperma (Kindb.) F.Schmidt (1868)
Spergularia longicaulis Pomel (1874)
Spergularia marina f. longicaulis (Pomel) Batt. (1888)
Spergularia marina var. asiatica (H.Hara) H.Hara (1941)
Spergularia marina var. cleistogama Y.X.Ma & D.L.Cui (2012)
Spergularia marina var. densiflora Hosny ex Hosni (1988)
Spergularia marina var. heterosperma (Fenzl) Neilr. (1859)
Spergularia marina var. melanocaulos (Merino) Merino (1897)
Spergularia marina var. simonii O.Deg. & I.Deg. (1962)
Spergularia marina var. stenopetala A.Chev. (1905)
Spergularia marina var. tenuis (Greene) R.Rossbach (1940)
Spergularia marina var. urbica (Leffler) Cout. (1913)
Spergularia media var. heterosperma Fenzl (1843)
Spergularia melanocaulos Merino (1895)
Spergularia miquelonensis Lebel (1868)
Spergularia modesta proles marina (L.) Samp. (1909)
Spergularia modesta proles urbica (Leffler) Samp. (1909)
Spergularia neglecta Syme (1864)
Spergularia pinguis Rouy & Foucaud (1896)
Spergularia rodrigoi Sennen & Mauricio (1936)
Spergularia rubra subsp. salina (J.Presl & C.Presl) Čelak. (1870)
Spergularia rubra var. dillenii (Lebel) Knoche (1921)
Spergularia rubra var. marina (L.) A.Gray (1848)
Spergularia rubra var. pinguis Fenzl (1843)
Spergularia rubra var. sperguloides (Lehm.) Ball (1877)
Spergularia salina (Pers.) J. Presl & C.Presl (1819)
Spergularia salina var. asiatica H.Hara (1937)
Spergularia salina var. australis (Lebel) Gürke (1899)
Spergularia salina var. involucrata (B.L.Rob.) Jeps. (1911)
Spergularia salina var. minor (S.Watson) B.L.Rob. (1894)
Spergularia salina var. sordida (Greene) Jeps. (1915)
Spergularia salina var. tenuior Rupr. (1860)
Spergularia salina var. tenuis (Greene) Jeps. (1911)
Spergularia salina var. urbica (Leffler) Gürke (1899)
Spergularia seminulifera Hy (1913)
Spergularia sparsiflora (Greene) A.Nelson (1909)
Spergularia sperguloides (Lehm.) Heynh. (1847)
Spergularia tenuis (Greene) B.L.Rob. (1894)
Spergularia tenuis var. involucrata B.L.Rob. (1897)
Spergularia uliginosa Pomel (1874)
Spergularia urbica Nyman (1878)
Stipularia marina (L.) Haw. (1812)
Tissa coreana (H.Lév.) Nakai (1934)
Tissa fauriei Nakai (1913)
Tissa leiosperma (Kindb.) A.Heller (1910)
Tissa marina (L.) Britton (1889)
Tissa marina var. minor (S.Watson) Britton (1901)
Tissa salina (J.Presl & C.Presl) Britton (1889)
Tissa salina var. involucrata (B.L.Rob.) Jeps. (1901)
Tissa salina var. sanfordii Greene (1891)
Tissa salina var. sordida Greene (1891)
Tissa salina var. tenuis (Greene) Jeps. (1901)
Tissa sparsiflora Greene (1895)
Tissa tenuis (Greene) Greene (1889)

Uses and Culture

USES

Natural History

Statewide Status

Naturalized

Island Status

Mokupāpapa (Kure atoll) Naturalized
Kuaihelani (Midway Atoll) Naturalized
Kaua'i Naturalized
O'ahu Naturalized
Molokai Naturalized
Lalo (French Frigate Shoals) Naturalized
Maui Naturalized

Dispersal Agents


Pollinators

Notes

  • Apparently used by Hawaiians medicinally. St. John (1973a) lisrs s. media (L.) K. Presl as adventive in Hawai'i; this appears to have been based on 3 specimens of S. marina at BISH from Koko Head misidentified as S. media by C. N. Forbes.
  • Description digitized by Mercedez
  • Description digitized from the Manual of the Flowering Plants of Hawaii
  • First collected on O'ahu in 1909

Bibliography

Name Published In: Enum. Pl.: 97 (1822)

Other References

Wagner et al. 1990:526 (Ku, Mi, FF, K, O, Mo, M)

Occurrences

SNo. Scientific Name Scientific Name Authorship Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Spergularia marina (L.) Besser summit trail s of kealia bare rocky clay of wide path PRESERVED_SPECIMEN degener, o. hatheway, w.h. collector number: 20670 Oahu BISH 5/14/1950
2 Spergularia marina (L.) Besser mokuleia beach, waialua Seasonally flooded bulldozed sand pit near ocean. PRESERVED_SPECIMEN degener, o. degener, i. collector number: 27846 Oahu BISH 2/1/1962
3 Spergularia marina (L.) Besser kamilonui valley, submerged in water by kuapa pond; bare soil; below kiawe zone. (near koko head.) PRESERVED_SPECIMEN egler, f.e. collector number: 37-110 Oahu BISH 7/11/1937
4 Spergularia marina (L.) Besser KAUAI: Salt Pond County Park. Halophytic vegetation of Batis maritima and Pluchea indica around salt pans. PRESERVED_SPECIMEN David H. Lorence 7438 Kauai PTBG 2/5/1994
5 Spergularia marina (L.) Besser kuaui. salt pond park west of burns field, hanapepe. PRESERVED_SPECIMEN t. flynn 1605 NY 3/18/1986
6 Spergularia marina (L.) Besser Salt Pond Park west of Burns Field, Hanapepe. PRESERVED_SPECIMEN Tim Flynn 1605 Kauai PTBG 3/18/1986
7 Spergularia marina (L.) Besser Salt Pond Beach Park area, ancient salt pans and littoral vegetation, western end of Hanapepe makai of Burns Field perimeter, shrubland/grassland/herbland. PRESERVED_SPECIMEN K. R. Wood 11131 Kauai PTBG 2/9/2005
8 Spergularia marina (L.) Besser Waimea Dist., Salt Pond County Park PRESERVED_SPECIMEN Lorence, D. H. 7438 Kauai US 2/5/1994
9 Spergularia marina (L.) Besser MAUI: E. Maui, Kanaha Pond Wildlife Sanctuary. Growing in sandy soil near observation kiosk at Hana Hwy. PRESERVED_SPECIMEN Hank Oppenheimer H100052 Maui PTBG 10/30/2000
10 Spergularia marina (L.) Besser Koko Head. Tuff sea cliffs opposite the Koko Head Rifle Range. PRESERVED_SPECIMEN Nagata, K. M. 1232 Oahu US 7/25/1974
11 Spergularia marina (L.) Besser tern islet, along southeastern edge of the runway. PRESERVED_SPECIMEN d. r. herbst 6247 French Frigate Shoals US 9/11/1978
12 Spergularia marina (L.) Besser kamilonui valley, by kuapa pond, below kiawe PRESERVED_SPECIMEN f. egler 37-110 Oahu US 7/11/1937
13 Spergularia marina (L.) Besser From the south side of the runway PRESERVED_SPECIMEN Long, C. R. 2247 Kure Atoll US 9/14/1964
14 Spergularia marina (L.) Besser Honolulu, north of Ala Moana Park, in sand deredged from Kewalo Basin PRESERVED_SPECIMEN Judd, A. F. s.n. Oahu US 5/25/1937
15 Spergularia marina (L.) Besser Near air field, Puolo, Hanapepe PRESERVED_SPECIMEN Degener, O.; Degener, I. 27353 Kauai US 3/14/1961
16 Spergularia marina (L.) Besser Barbers Point, coral plain, near beach. PRESERVED_SPECIMEN Ning, W. S. s.n. Oahu US 8/8/1937
17 Spergularia marina (L.) Besser along Kaahumanu Highway east of Waikoloa Junction. PRESERVED_SPECIMEN Herbst, D. R.; Ishikawa, S. 5386 Hawaii US 6/24/1975
18 Spergularia marina (L.) Besser hamakua marsh, kailua, mauka (mountainside) of hamakua drive PRESERVED_SPECIMEN c. imada, a. engilis & a. yuen 92-6 Oahu US 4/4/1992
19 Spergularia marina (L.) Besser waianae. PRESERVED_SPECIMEN f. r. fosberg & v. o. fosberg 13624 Oahu US 3/21/1937
20 Spergularia marina (L.) Besser mokuleia PRESERVED_SPECIMEN o. degener 17160 Oahu US 3/14/1933
21 Spergularia marina (L.) Besser Koko Head. Tuff sea cliffs opposite the Koko Head Rifle Range PRESERVED_SPECIMEN Nagata, K. M. 1230 Oahu US 7/25/1974
22 Spergularia marina (L.) Besser koko head. tuff sea cliffs opposite the koko head rifle range. PRESERVED_SPECIMEN k. nagata 1232 Oahu US 7/25/1974
23 Spergularia marina (L.) Besser Kamilonui Valley, by Kuapa Pond, below Kiawe PRESERVED_SPECIMEN Egler, F. E. 37-110 Oahu US 7/11/1937
24 Spergularia marina (L.) Besser along kaahumanu highway east of waikoloa junction. PRESERVED_SPECIMEN d. r. herbst & s. ishikawa 5386 Hawaii US 6/24/1975
25 Spergularia marina (L.) Besser near air field, puolo, hanapepe PRESERVED_SPECIMEN o. degener & i. degener 27353 Kauai US 3/14/1961
26 Spergularia marina (L.) Besser koko head. tuff sea cliffs opposite the koko head rifle range PRESERVED_SPECIMEN k. nagata 1230 Maui US 7/25/1974
27 Spergularia marina (L.) Besser waimea dist., salt pond county park PRESERVED_SPECIMEN d. h. lorence 7438 Kauai US 2/5/1994
28 Spergularia marina (L.) Besser Hamakua Marsh, Kailua, mauka (mountainside) of Hamakua Drive PRESERVED_SPECIMEN Imada, C. T.; Engilis, A.; Yuen, A. 92-6 Oahu US 4/4/1992
29 Spergularia marina (L.) Besser barbers point, coral plain, near beach. PRESERVED_SPECIMEN w. ning s.n. Oahu US 8/8/1937
30 Spergularia marina (L.) Besser Pouhala Marsh, Waipahu. PRESERVED_SPECIMEN Crago, L. M.; Erickson, T. 2005-024 Oahu US 1/24/2005
31 Spergularia marina (L.) Besser from the south side of the runway PRESERVED_SPECIMEN c. long 2247 Kure Atoll US 9/14/1964
32 Spergularia marina (L.) Besser Honolulu, North of Ala Moana Park, in sand dredged from Kewalo Basin PRESERVED_SPECIMEN Judd Jr., A.F. s.n. Oahu BISH 5/25/1937
33 Spergularia marina (L.) Besser kapuaina coconut grove open ground near beach PRESERVED_SPECIMEN pekelo jr., n. collector number: 35 Molokai BISH 3/8/1974
34 Spergularia marina (L.) Besser near koko hd. on the beach PRESERVED_SPECIMEN forbes, c.n. collector number: 1913.0 Oahu BISH 7/1/1914
35 Spergularia marina (L.) Besser Kapuaiwa Coconut Grove open ground near beach PRESERVED_SPECIMEN Pekelo Jr., N. 35 Molokai BISH 3/8/1974
36 Spergularia marina (L.) Besser Mokuleia Beach, Waialua, sand pit near ocean PRESERVED_SPECIMEN Degener, O.; Degener, I. 27846 Oahu US 2/2/1962
37 Spergularia marina (L.) Besser eastern island, edge of runway PRESERVED_SPECIMEN herbst, d.r. collector number: 6409 Oahu BISH 6/30/1980
38 Spergularia marina (L.) Besser sandspit in front of kuapa pond between kuliouou and portlock In sand just above high water mark PRESERVED_SPECIMEN herbst, d.r. collector number: 413 Oahu BISH 4/5/1967
39 Spergularia marina (L.) Besser honolulu, kapiolani park PRESERVED_SPECIMEN munro, g.c. collector number: 58 Oahu BISH 4/14/1950
40 Spergularia marina (L.) Besser ewa, barbers point, on dry open flat near beach PRESERVED_SPECIMEN hosaka, e.y. collector number: 1352 Oahu BISH 3/22/1936
41 Spergularia marina (L.) Besser honolulu, north of ala moana park, in sand deredged from kewalo basin PRESERVED_SPECIMEN a. judd s.n. Oahu US 5/25/1937
42 Spergularia marina (L.) Besser Mokuleia PRESERVED_SPECIMEN Degener, O. 17160 Oahu US 3/14/1933
43 Spergularia marina (L.) Besser E side of Hale o Lono Harbor In sandy soil. PRESERVED_SPECIMEN Oppenheimer, H.L. H50629 Molokai BISH 5/15/2005
44 Spergularia marina (L.) Besser mokuleia beach, waialua, sand pit near ocean PRESERVED_SPECIMEN o. degener & i. degener 27846 Oahu US 2/2/1962
45 Spergularia marina (L.) Besser Koko Head Oahu PRESERVED_SPECIMEN Forbes, C.N. 1448 o. Oahu BISH 6/11/1909
46 Spergularia marina (L.) Besser Waianae. PRESERVED_SPECIMEN Fosberg, F. R.; Fosberg, V. O. 13624 Oahu US 3/21/1937
47 Spergularia marina (L.) Besser ewa beach PRESERVED_SPECIMEN bennett, t. collector number: 174 Maui BISH 2/24/1982
48 Spergularia marina (L.) Besser pouhala marsh, waipahu. PRESERVED_SPECIMEN l. crago & t. erickson 2005-024 Oahu US 1/24/2005
49 Spergularia marina (L.) Besser roadside near niu PRESERVED_SPECIMEN forbes, c.n. collector number: 2422.0 Oahu BISH 2/16/1917
50 Spergularia marina (L.) Besser Tern Islet, along southeastern edge of the runway. PRESERVED_SPECIMEN Herbst, D. R. 6247 French Frigate Shoals US 9/11/1978
51 Spergularia marina (L.) Besser damp spot, waipio Ferns in 6" pots inside plastic-roofed house. PRESERVED_SPECIMEN pearsall, g.a. Hawaii BISH 3/19/1954
52 Spergularia marina (L.) Besser makai of aquaculture site of kahuku Collected in solid, dwarfed strand community of Heliotropium anomalum, Jacquemontia, Fimbristylis, Nama, Euphorbia degeneri and Lycium. PRESERVED_SPECIMEN stemmermann, r.l. palmer, d.; obata, j. collector number: 3703 Oahu BISH 2/11/1979
53 Spergularia marina (L.) Besser Near Koko Head on the beach with J.F.S.Stokes PRESERVED_SPECIMEN Forbes, C.N. 1913.O Oahu BISH 7/1/1914
54 Spergularia marina (L.) Besser among branches of atriplex sp. golden crown beard, mixed grasses, and batis sp. west of barbers point deep draft harbor Plants were seeding and the population was robust. PRESERVED_SPECIMEN funk, e.j. lee, m.; b.ching collector number: 403 Oahu BISH 5/1/1990
55 Spergularia marina (L.) Besser Roadside near Niu PRESERVED_SPECIMEN Forbes, C.N. 2422.O Oahu BISH 2/16/1917
56 Spergularia marina (L.) Besser Tern Islet PRESERVED_SPECIMEN Lamoureux, C.H. 1663 French Frigate Shoals BISH 9/2/1961
57 Spergularia marina (L.) Besser Waialua, Kawailua, roadside PRESERVED_SPECIMEN Bush, W. s.n. Oahu BISH 3/5/1933
58 Spergularia marina (L.) Besser koko head. tuff sea cliffs opposite the koko head rifle range PRESERVED_SPECIMEN nagata, k.m. collector number: 1230 Oahu BISH 7/25/1974
59 Spergularia marina (L.) Besser Kahului, near brackish marsh PRESERVED_SPECIMEN Degener, O. 35810 Maui BISH 7/11/1927
60 Spergularia marina (L.) Besser salt pond park w of burns field, hanapepe, around pavillion Moist woods PRESERVED_SPECIMEN flynn, t. collector number: 1605 Molokai BISH 3/18/1986
61 Spergularia marina (L.) Besser waianae mts., piko trail PRESERVED_SPECIMEN kerr, m. collector number: 38-0 Oahu BISH 5/5/1946
62 Spergularia marina (L.) Besser Pouhala Marsh, Waipahu Growing at edge and higher up on the bank of Pouhala. In wet to dry soil within the saltlines. Large patches in full sun to medium shade. PRESERVED_SPECIMEN Crago, L.M. 2005-024 Oahu BISH 1/24/2005
63 Spergularia marina (L.) Besser Honolulu, Kapiolani Park PRESERVED_SPECIMEN Munro, G.C. 58 Oahu BISH 4/14/1950
64 Spergularia marina (L.) Besser Koko Head. Tuff sea cliffs opposite the Koko Head Rifle Range In small communities in exposed solution basins. Associated species: Heliotropium, Lipochaeta, Batis. PRESERVED_SPECIMEN Nagata, K.M. 1232 Oahu BISH 7/25/1974
65 Spergularia marina (L.) Besser kahului, near brackish marsh PRESERVED_SPECIMEN degener, o. collector number: 35810 Maui BISH 7/11/1927
66 Spergularia marina (L.) Besser ewa plains PRESERVED_SPECIMEN higashino, p.k. collector number: 8189 Kauai BISH 2/11/1979
67 Spergularia marina (L.) Besser S. of Ewa, 1 mile W of Oneula, in salt flats back of beach Weed PRESERVED_SPECIMEN St. John, H. 24804 Oahu BISH 4/27/1952
68 Spergularia marina (L.) Besser Waianae Mts., Piko trail PRESERVED_SPECIMEN Kerr, M. 38-0 Oahu BISH 5/5/1946
69 Spergularia marina (L.) Besser E side of Hale o Lono Harbor In sandy soil. PRESERVED_SPECIMEN Oppenheimer, H.L. Tangalin, N.; Perlman, S.; Garnett, B. Collector Number: H50629 Molokai BISH 5/15/2005
70 Spergularia marina (L.) Besser Ewa, Barbers Point, on dry open flat near beach PRESERVED_SPECIMEN Hosaka, E.Y. 1352 Oahu BISH 3/22/1936
71 Spergularia marina (L.) Besser beach near koko head PRESERVED_SPECIMEN forbes, c.n. collector number: 1447 Oahu BISH 2/15/1909
72 Spergularia marina (L.) Besser Barbers Point, east of the lighthouse on dry thin soil back of the beach in the open PRESERVED_SPECIMEN Egler, F.E. 37-274 Oahu BISH 8/11/1937
73 Spergularia marina (L.) Besser Salt Pond Park W of Burns Field, Hanapepe, around pavillion lawn PRESERVED_SPECIMEN Flynn, T. 1605 Kauai BISH 3/18/1986
74 Spergularia marina (L.) Besser Kure Atoll PRESERVED_SPECIMEN Lamoureux, C.H. 1910 Kure Atoll BISH 9/13/1961
75 Spergularia marina (L.) Besser honolulu, oahu. herb medicines, native name mimi ilio, (urine dog) PRESERVED_SPECIMEN pukui, m.k. Oahu BISH 8/8/1931
76 Spergularia marina (L.) Besser Summit trail south of Kealia, Oahu. Growing only on bare rocky clay of wide path PRESERVED_SPECIMEN Degener, O. 20670 Oahu BISH 3/14/1950
77 Spergularia marina (L.) Besser waimea, salt pond county park, w side of salt pan and adjacent edge of park Halephytic vegetation of Batis maritima and Pluchea indica PRESERVED_SPECIMEN lorence, d.h. collector number: 7438 Kauai BISH 2/5/1994
78 Spergularia marina (L.) Besser beach e of barber's point PRESERVED_SPECIMEN fosberg, f.r. collector number: 29528 Molokai BISH 2/4/1948
79 Spergularia marina (L.) Besser tern islet PRESERVED_SPECIMEN lamoureux, c.h. collector number: 1663 French Frigate Shoals BISH 9/2/1961
80 Spergularia marina (L.) Besser Eastern Island, edge of runway PRESERVED_SPECIMEN Herbst, D.R. 6409 Midway Atoll BISH 6/30/1980
81 Spergularia marina (L.) Besser Tern Islet, SE edge of runway PRESERVED_SPECIMEN Herbst, D.R. 6247 French Frigate Shoals BISH 9/11/1978
82 Spergularia marina (L.) Besser Summit trail S of Kealia bare rocky clay of wide path PRESERVED_SPECIMEN Degener, O. 20670 Oahu BISH 5/14/1950
83 Spergularia marina (L.) Besser summit trail south of kealia, oahu. growing only on bare rocky clay of wide path PRESERVED_SPECIMEN degener, o. collector number: 20670 Oahu BISH 3/14/1950
84 Spergularia marina (L.) Besser Beach near Koko Head PRESERVED_SPECIMEN Forbes, C.N. 1447 Oahu BISH 2/15/1909
85 Spergularia marina (L.) Besser Makai of aquaculture site of Kahuku Collected in solid, dwarfed strand community of Heliotropium anomalum, Jacquemontia, Fimbristylis, Nama, Euphorbia degeneri and Lycium. PRESERVED_SPECIMEN Stemmermann, R.L. 3703 Oahu BISH 2/11/1979
86 Spergularia marina (L.) Besser hamakua marsh, kailua, mauka (mountainside) of hamakua drive Low groundcover growing on dried mudflats with Bacopa, Paspalum. PRESERVED_SPECIMEN imada, c.t. engilis, a.; yuen, a.; dr.duk collector number: 92-6 Oahu BISH 4/4/1992
87 Spergularia marina (L.) Besser Sand Island, SE end of inner harbor mudflat PreservedSpecimen Herbst, D.R. Takeuchi, W.N. Collector Number: 6349 Midway Atoll BISH Specimen 6/29/1980
88 Spergularia marina (L.) Besser Eastern Island, edge of runway PreservedSpecimen Herbst, D.R. Collector Number: 6409 Midway Atoll BISH Specimen 6/30/1980
89 Spergularia marina (L.) Besser beach E of Barber's Point along shaded path in Prosopis on coral limestone PRESERVED_SPECIMEN Fosberg, F.R. 29528 Oahu BISH 2/4/1948
90 Spergularia marina (L.) Besser Ewa. Pohakupuna Rd PRESERVED_SPECIMEN Woolford, H.A. s.n. Oahu BISH
91 Spergularia marina (L.) Besser Near Koko Hd. on the beach PRESERVED_SPECIMEN Forbes, C.N. 1913.O Oahu BISH 7/1/1914
92 Spergularia marina (L.) Besser tern islet, se edge of runway PRESERVED_SPECIMEN herbst, d.r. collector number: 6247 French Frigate Shoals BISH 9/11/1978
93 Spergularia marina (L.) Besser Hamakua Marsh, Kailua, mauka (mountainside) of Hamakua Drive Low groundcover growing on dried mudflats with Bacopa, Paspalum. PRESERVED_SPECIMEN Imada, C.T. 92-6 Oahu BISH 4/4/1992
94 Spergularia marina (L.) Besser Waipio Peninsula; Waipio, Pouhala Marsh between Kapakahi & Waikele Streams Occasional herb on mudflats adjacent to Batis marshland & open water. PRESERVED_SPECIMEN Imada, C.T. 96-10 Oahu BISH 12/18/1996
95 Spergularia marina (L.) Besser s. of ewa, 1 mile w of oneula, in salt flats back of beach Weed PRESERVED_SPECIMEN st.john, h. collector number: 24804 Oahu BISH 4/27/1952
96 Spergularia marina (L.) Besser honolulu, north of ala moana park, in sand dredged from kewalo basin PRESERVED_SPECIMEN judd jr., a.f. Oahu BISH 5/25/1937
97 Spergularia marina (L.) Besser Kamilonui Valley, submerged in water by Kuapa Pond; bare soil; below Kiawe zone. (Near Koko Head.) PRESERVED_SPECIMEN Egler, F.E. 37-110 Oahu BISH 7/11/1937
98 Spergularia marina (L.) Besser near koko head. fish ponds outer brook PRESERVED_SPECIMEN bennett, t. collector number: 1501 Oahu BISH 6/11/1909
99 Spergularia marina (L.) Besser sand island, se end of inner harbor In dense rich forest. PRESERVED_SPECIMEN herbst, d.r. takeuchi, w.n. collector number: 6349 Hawaii BISH 6/29/1980
100 Spergularia marina (L.) Besser near haleiwa. marshy land PRESERVED_SPECIMEN hilliaurs Oahu BISH 3/17/1930
101 Spergularia marina (L.) Besser koko head oahu PRESERVED_SPECIMEN forbes, c.n. collector number: 1448 o. Oahu BISH 6/11/1909
102 Spergularia marina (L.) Besser Mokuleia Beach, Waialua Seasonally flooded bulldozed sand pit near ocean. PRESERVED_SPECIMEN Degener, O. 27846 Oahu BISH 2/1/1962
103 Spergularia marina (L.) Besser Pouhala Marsh, Waipahu Growing at edge and higher up on the bank of Pouhala. In wet to dry soil within the saltlines. Large patches in full sun to medium shade. PRESERVED_SPECIMEN Crago, L.M. Erickson, T. Collector Number: 2005-024 Oahu BISH 1/24/2005
104 Spergularia marina (L.) Besser e side of hale o lono harbor In sandy soil. PRESERVED_SPECIMEN oppenheimer, h.l. tangalin, n.; perlman, s.; garnett, b. collector number: h50629 Molokai BISH 5/15/2005
105 Spergularia marina (L.) Besser Near Haleiwa. Marshy land PRESERVED_SPECIMEN Hilliaurs s.n. Oahu BISH 3/17/1930
106 Spergularia marina (L.) Besser koko head. tuff sea cliffs opposite the koko head rifle range In small communities in exposed solution basins. Associated species: Heliotropium, Lipochaeta, Batis. PRESERVED_SPECIMEN nagata, k.m. collector number: 1232 Oahu BISH 7/25/1974
107 Spergularia marina (L.) Besser ewa, pohakupuna rd PRESERVED_SPECIMEN woolford, h.a. Oahu BISH 2/10/1957
108 Spergularia marina (L.) Besser near air field, puolo, hanapepe PRESERVED_SPECIMEN degener, o. degener, i. collector number: 27353 Kauai BISH 3/14/1961
109 Spergularia marina (L.) Besser barbers point, east of the lighthouse on dry thin soil back of the beach in the open PRESERVED_SPECIMEN egler, f.e. collector number: 37-274 Oahu BISH 8/11/1937
110 Spergularia marina (L.) Besser Near Koko Head. Fish Ponds outer brook PRESERVED_SPECIMEN Bennett, T. 1501 Oahu BISH 6/11/1909
111 Spergularia marina (L.) Besser Damp spot, Waipio PRESERVED_SPECIMEN Pearsall, G.A. s.n. Oahu BISH 3/19/1954
112 Spergularia marina (L.) Besser ewa, pohakupuna rd PRESERVED_SPECIMEN woolford, h.a. collector number: s.n. Oahu BISH 2/10/1957
113 Spergularia marina (L.) Besser near air field, Puolo, Hanapepe naturalized in mud flats PRESERVED_SPECIMEN Degener, O. 27353 Kauai BISH 3/14/1961
114 Spergularia marina (L.) Besser Ewa Plains Prosopis, Leucaena, Cynodon, Setaria, Many introduced plants. PRESERVED_SPECIMEN Higashino, P.K. 8189 Oahu BISH 2/11/1979
115 Spergularia marina (L.) Besser waipio peninsula; waipio, pouhala marsh between kapakahi & waikele streams Occasional herb on mudflats adjacent to Batis marshland & open water. PRESERVED_SPECIMEN imada, c.t. nishida, g. collector number: 96-10 Oahu BISH 12/18/1996
116 Spergularia marina (L.) Besser Koko Head. Tuff sea cliffs opposite the Koko Head Rifle Range In small but dense communities in solution basins. Associated with Batis, Heliotropium, Lipochaeta. PRESERVED_SPECIMEN Nagata, K.M. 1230 Oahu BISH 7/25/1974
117 Spergularia marina (L.) Besser Waimea, Salt Pond County Park, W side of salt pan and adjacent edge of park Halephytic vegetation of Batis maritima and Pluchea indica PRESERVED_SPECIMEN Lorence, D.H. 7438 Kauai BISH 2/5/1994
118 Spergularia marina (L.) Besser Sand Island, SE end of inner harbor mudflat PRESERVED_SPECIMEN Herbst, D.R. 6349 Midway Atoll BISH 6/29/1980
119 Spergularia marina (L.) Besser Sandspit in front of Kuapa Pond between Kuliouou and Portlock In sand just above high water mark PRESERVED_SPECIMEN Herbst, D.R. 413 Oahu BISH 4/5/1967
120 Spergularia marina (L.) Besser waialua, kawailua, roadside PRESERVED_SPECIMEN bush, w. topping, d.l. collector number: s.n. Oahu BISH 3/5/1933
121 Spergularia marina (L.) Besser Honolulu, Oahu. Herb medicines, native name MIMI ILIO, (urine dog) PRESERVED_SPECIMEN Pukui, M.K. s.n. Oahu BISH 4/14/1905
122 Spergularia marina (L.) Besser Ewa Beach PRESERVED_SPECIMEN Bennett, T. 174 Oahu BISH 2/24/1982
123 Spergularia marina (L.) Besser among branches of Atriplex sp. golden crown beard, mixed grasses, and Batis sp. west of Barbers Point Deep Draft harbor Plants were seeding and the population was robust. PRESERVED_SPECIMEN Funk, E.J. 403 Oahu BISH 5/1/1990
124 Spergularia marina (L.) Besser near koko head on the beach with j.f.s.stokes PRESERVED_SPECIMEN forbes, c.n. collector number: 1913.6 Oahu BISH 7/1/1914
125 Spergularia marina (L.) Besser pouhala marsh, waipahu Growing at edge and higher up on the bank of Pouhala. In wet to dry soil within the saltlines. Large patches in full sun to medium shade. PRESERVED_SPECIMEN crago, l.m. erickson, t. collector number: 2005-024 Oahu BISH 1/24/2005
126 Spergularia marina (L.) Besser Ca. 200 yds, east of the light house at Barber's Point. Arid, sandy region behind beach. Assoc. w/ Pluchea indica, P. odorata, Batis, Lycium, Sesuvium, Spergularia, sporobolus virg. Gregarious, naturalized. PRESERVED_SPECIMEN Herbst, D.R. 505 Oahu BISH 5/21/1967
127 Spergularia marina (L.) Besser kure atoll PRESERVED_SPECIMEN lamoureux, c.h. collector number: 1910 Kure Atoll BISH 9/13/1961
128 Spergularia marina (L.) Besser ewa. pohakupuna rd PRESERVED_SPECIMEN woolford, h.a. Oahu BISH
129 Spergularia marina (L.) Besser summit trail s of kealia bare rocky clay of wide path PRESERVED_SPECIMEN degener, o. hatheway, w.h. collector number: 20670 Oahu BISH 5/14/1950
130 Spergularia marina (L.) Besser mokuleia beach, waialua Seasonally flooded bulldozed sand pit near ocean. PRESERVED_SPECIMEN degener, o. degener, i. collector number: 27846 Oahu BISH 2/1/1962
131 Spergularia marina (L.) Besser kamilonui valley, submerged in water by kuapa pond; bare soil; below kiawe zone. (near koko head.) PRESERVED_SPECIMEN egler, f.e. collector number: 37-110 Oahu BISH 7/11/1937
132 Spergularia marina (L.) Besser KAUAI: Salt Pond County Park. Halophytic vegetation of Batis maritima and Pluchea indica around salt pans. PRESERVED_SPECIMEN David H. Lorence 7438 Kauai PTBG 2/5/1994
133 Spergularia marina (L.) Besser kuaui. salt pond park west of burns field, hanapepe. PRESERVED_SPECIMEN t. flynn 1605 NY 3/18/1986
134 Spergularia marina (L.) Besser Salt Pond Park west of Burns Field, Hanapepe. PRESERVED_SPECIMEN Tim Flynn 1605 Kauai PTBG 3/18/1986
135 Spergularia marina (L.) Besser Salt Pond Beach Park area, ancient salt pans and littoral vegetation, western end of Hanapepe makai of Burns Field perimeter, shrubland/grassland/herbland. PRESERVED_SPECIMEN K. R. Wood 11131 Kauai PTBG 2/9/2005
136 Spergularia marina (L.) Besser Waimea Dist., Salt Pond County Park PRESERVED_SPECIMEN Lorence, D. H. 7438 Kauai US 2/5/1994
137 Spergularia marina (L.) Besser MAUI: E. Maui, Kanaha Pond Wildlife Sanctuary. Growing in sandy soil near observation kiosk at Hana Hwy. PRESERVED_SPECIMEN Hank Oppenheimer H100052 Maui PTBG 10/30/2000
138 Spergularia marina (L.) Besser Koko Head. Tuff sea cliffs opposite the Koko Head Rifle Range. PRESERVED_SPECIMEN Nagata, K. M. 1232 Oahu US 7/25/1974
139 Spergularia marina (L.) Besser tern islet, along southeastern edge of the runway. PRESERVED_SPECIMEN d. r. herbst 6247 French Frigate Shoals US 9/11/1978
140 Spergularia marina (L.) Besser kamilonui valley, by kuapa pond, below kiawe PRESERVED_SPECIMEN f. egler 37-110 Oahu US 7/11/1937
141 Spergularia marina (L.) Besser From the south side of the runway PRESERVED_SPECIMEN Long, C. R. 2247 Kure Atoll US 9/14/1964
142 Spergularia marina (L.) Besser Honolulu, north of Ala Moana Park, in sand deredged from Kewalo Basin PRESERVED_SPECIMEN Judd, A. F. s.n. Oahu US 5/25/1937
143 Spergularia marina (L.) Besser Near air field, Puolo, Hanapepe PRESERVED_SPECIMEN Degener, O.; Degener, I. 27353 Kauai US 3/14/1961
144 Spergularia marina (L.) Besser Barbers Point, coral plain, near beach. PRESERVED_SPECIMEN Ning, W. S. s.n. Oahu US 8/8/1937
145 Spergularia marina (L.) Besser along Kaahumanu Highway east of Waikoloa Junction. PRESERVED_SPECIMEN Herbst, D. R.; Ishikawa, S. 5386 Hawaii US 6/24/1975
146 Spergularia marina (L.) Besser hamakua marsh, kailua, mauka (mountainside) of hamakua drive PRESERVED_SPECIMEN c. imada, a. engilis & a. yuen 92-6 Oahu US 4/4/1992
147 Spergularia marina (L.) Besser waianae. PRESERVED_SPECIMEN f. r. fosberg & v. o. fosberg 13624 Oahu US 3/21/1937
148 Spergularia marina (L.) Besser mokuleia PRESERVED_SPECIMEN o. degener 17160 Oahu US 3/14/1933
149 Spergularia marina (L.) Besser Koko Head. Tuff sea cliffs opposite the Koko Head Rifle Range PRESERVED_SPECIMEN Nagata, K. M. 1230 Oahu US 7/25/1974
150 Spergularia marina (L.) Besser koko head. tuff sea cliffs opposite the koko head rifle range. PRESERVED_SPECIMEN k. nagata 1232 Oahu US 7/25/1974
151 Spergularia marina (L.) Besser Kamilonui Valley, by Kuapa Pond, below Kiawe PRESERVED_SPECIMEN Egler, F. E. 37-110 Oahu US 7/11/1937
152 Spergularia marina (L.) Besser along kaahumanu highway east of waikoloa junction. PRESERVED_SPECIMEN d. r. herbst & s. ishikawa 5386 Hawaii US 6/24/1975
153 Spergularia marina (L.) Besser near air field, puolo, hanapepe PRESERVED_SPECIMEN o. degener & i. degener 27353 Kauai US 3/14/1961
154 Spergularia marina (L.) Besser koko head. tuff sea cliffs opposite the koko head rifle range PRESERVED_SPECIMEN k. nagata 1230 Maui US 7/25/1974
155 Spergularia marina (L.) Besser waimea dist., salt pond county park PRESERVED_SPECIMEN d. h. lorence 7438 Kauai US 2/5/1994
156 Spergularia marina (L.) Besser Hamakua Marsh, Kailua, mauka (mountainside) of Hamakua Drive PRESERVED_SPECIMEN Imada, C. T.; Engilis, A.; Yuen, A. 92-6 Oahu US 4/4/1992
157 Spergularia marina (L.) Besser barbers point, coral plain, near beach. PRESERVED_SPECIMEN w. ning s.n. Oahu US 8/8/1937
158 Spergularia marina (L.) Besser Pouhala Marsh, Waipahu. PRESERVED_SPECIMEN Crago, L. M.; Erickson, T. 2005-024 Oahu US 1/24/2005
159 Spergularia marina (L.) Besser from the south side of the runway PRESERVED_SPECIMEN c. long 2247 Kure Atoll US 9/14/1964
160 Spergularia marina (L.) Besser Honolulu, North of Ala Moana Park, in sand dredged from Kewalo Basin PRESERVED_SPECIMEN Judd Jr., A.F. s.n. Oahu BISH 5/25/1937
161 Spergularia marina (L.) Besser kapuaina coconut grove open ground near beach PRESERVED_SPECIMEN pekelo jr., n. collector number: 35 Molokai BISH 3/8/1974
162 Spergularia marina (L.) Besser near koko hd. on the beach PRESERVED_SPECIMEN forbes, c.n. collector number: 1913.0 Oahu BISH 7/1/1914
163 Spergularia marina (L.) Besser Kapuaiwa Coconut Grove open ground near beach PRESERVED_SPECIMEN Pekelo Jr., N. 35 Molokai BISH 3/8/1974
164 Spergularia marina (L.) Besser Mokuleia Beach, Waialua, sand pit near ocean PRESERVED_SPECIMEN Degener, O.; Degener, I. 27846 Oahu US 2/2/1962
165 Spergularia marina (L.) Besser eastern island, edge of runway PRESERVED_SPECIMEN herbst, d.r. collector number: 6409 Oahu BISH 6/30/1980
166 Spergularia marina (L.) Besser sandspit in front of kuapa pond between kuliouou and portlock In sand just above high water mark PRESERVED_SPECIMEN herbst, d.r. collector number: 413 Oahu BISH 4/5/1967
167 Spergularia marina (L.) Besser honolulu, kapiolani park PRESERVED_SPECIMEN munro, g.c. collector number: 58 Oahu BISH 4/14/1950
168 Spergularia marina (L.) Besser ewa, barbers point, on dry open flat near beach PRESERVED_SPECIMEN hosaka, e.y. collector number: 1352 Oahu BISH 3/22/1936
169 Spergularia marina (L.) Besser honolulu, north of ala moana park, in sand deredged from kewalo basin PRESERVED_SPECIMEN a. judd s.n. Oahu US 5/25/1937
170 Spergularia marina (L.) Besser Mokuleia PRESERVED_SPECIMEN Degener, O. 17160 Oahu US 3/14/1933
171 Spergularia marina (L.) Besser E side of Hale o Lono Harbor In sandy soil. PRESERVED_SPECIMEN Oppenheimer, H.L. H50629 Molokai BISH 5/15/2005
172 Spergularia marina (L.) Besser mokuleia beach, waialua, sand pit near ocean PRESERVED_SPECIMEN o. degener & i. degener 27846 Oahu US 2/2/1962
173 Spergularia marina (L.) Besser Koko Head Oahu PRESERVED_SPECIMEN Forbes, C.N. 1448 o. Oahu BISH 6/11/1909
174 Spergularia marina (L.) Besser Waianae. PRESERVED_SPECIMEN Fosberg, F. R.; Fosberg, V. O. 13624 Oahu US 3/21/1937
175 Spergularia marina (L.) Besser ewa beach PRESERVED_SPECIMEN bennett, t. collector number: 174 Maui BISH 2/24/1982
176 Spergularia marina (L.) Besser pouhala marsh, waipahu. PRESERVED_SPECIMEN l. crago & t. erickson 2005-024 Oahu US 1/24/2005
177 Spergularia marina (L.) Besser roadside near niu PRESERVED_SPECIMEN forbes, c.n. collector number: 2422.0 Oahu BISH 2/16/1917
178 Spergularia marina (L.) Besser Tern Islet, along southeastern edge of the runway. PRESERVED_SPECIMEN Herbst, D. R. 6247 French Frigate Shoals US 9/11/1978
179 Spergularia marina (L.) Besser damp spot, waipio Ferns in 6" pots inside plastic-roofed house. PRESERVED_SPECIMEN pearsall, g.a. Hawaii BISH 3/19/1954
180 Spergularia marina (L.) Besser makai of aquaculture site of kahuku Collected in solid, dwarfed strand community of Heliotropium anomalum, Jacquemontia, Fimbristylis, Nama, Euphorbia degeneri and Lycium. PRESERVED_SPECIMEN stemmermann, r.l. palmer, d.; obata, j. collector number: 3703 Oahu BISH 2/11/1979
181 Spergularia marina (L.) Besser Near Koko Head on the beach with J.F.S.Stokes PRESERVED_SPECIMEN Forbes, C.N. 1913.O Oahu BISH 7/1/1914
182 Spergularia marina (L.) Besser among branches of atriplex sp. golden crown beard, mixed grasses, and batis sp. west of barbers point deep draft harbor Plants were seeding and the population was robust. PRESERVED_SPECIMEN funk, e.j. lee, m.; b.ching collector number: 403 Oahu BISH 5/1/1990
183 Spergularia marina (L.) Besser Roadside near Niu PRESERVED_SPECIMEN Forbes, C.N. 2422.O Oahu BISH 2/16/1917
184 Spergularia marina (L.) Besser Tern Islet PRESERVED_SPECIMEN Lamoureux, C.H. 1663 French Frigate Shoals BISH 9/2/1961
185 Spergularia marina (L.) Besser Waialua, Kawailua, roadside PRESERVED_SPECIMEN Bush, W. s.n. Oahu BISH 3/5/1933
186 Spergularia marina (L.) Besser koko head. tuff sea cliffs opposite the koko head rifle range PRESERVED_SPECIMEN nagata, k.m. collector number: 1230 Oahu BISH 7/25/1974
187 Spergularia marina (L.) Besser Kahului, near brackish marsh PRESERVED_SPECIMEN Degener, O. 35810 Maui BISH 7/11/1927
188 Spergularia marina (L.) Besser salt pond park w of burns field, hanapepe, around pavillion Moist woods PRESERVED_SPECIMEN flynn, t. collector number: 1605 Molokai BISH 3/18/1986
189 Spergularia marina (L.) Besser waianae mts., piko trail PRESERVED_SPECIMEN kerr, m. collector number: 38-0 Oahu BISH 5/5/1946
190 Spergularia marina (L.) Besser Pouhala Marsh, Waipahu Growing at edge and higher up on the bank of Pouhala. In wet to dry soil within the saltlines. Large patches in full sun to medium shade. PRESERVED_SPECIMEN Crago, L.M. 2005-024 Oahu BISH 1/24/2005
191 Spergularia marina (L.) Besser Honolulu, Kapiolani Park PRESERVED_SPECIMEN Munro, G.C. 58 Oahu BISH 4/14/1950
192 Spergularia marina (L.) Besser Koko Head. Tuff sea cliffs opposite the Koko Head Rifle Range In small communities in exposed solution basins. Associated species: Heliotropium, Lipochaeta, Batis. PRESERVED_SPECIMEN Nagata, K.M. 1232 Oahu BISH 7/25/1974
193 Spergularia marina (L.) Besser kahului, near brackish marsh PRESERVED_SPECIMEN degener, o. collector number: 35810 Maui BISH 7/11/1927
194 Spergularia marina (L.) Besser ewa plains PRESERVED_SPECIMEN higashino, p.k. collector number: 8189 Kauai BISH 2/11/1979
195 Spergularia marina (L.) Besser S. of Ewa, 1 mile W of Oneula, in salt flats back of beach Weed PRESERVED_SPECIMEN St. John, H. 24804 Oahu BISH 4/27/1952
196 Spergularia marina (L.) Besser Waianae Mts., Piko trail PRESERVED_SPECIMEN Kerr, M. 38-0 Oahu BISH 5/5/1946
197 Spergularia marina (L.) Besser E side of Hale o Lono Harbor In sandy soil. PRESERVED_SPECIMEN Oppenheimer, H.L. Tangalin, N.; Perlman, S.; Garnett, B. Collector Number: H50629 Molokai BISH 5/15/2005
198 Spergularia marina (L.) Besser Ewa, Barbers Point, on dry open flat near beach PRESERVED_SPECIMEN Hosaka, E.Y. 1352 Oahu BISH 3/22/1936
199 Spergularia marina (L.) Besser beach near koko head PRESERVED_SPECIMEN forbes, c.n. collector number: 1447 Oahu BISH 2/15/1909
200 Spergularia marina (L.) Besser Barbers Point, east of the lighthouse on dry thin soil back of the beach in the open PRESERVED_SPECIMEN Egler, F.E. 37-274 Oahu BISH 8/11/1937
201 Spergularia marina (L.) Besser Salt Pond Park W of Burns Field, Hanapepe, around pavillion lawn PRESERVED_SPECIMEN Flynn, T. 1605 Kauai BISH 3/18/1986
202 Spergularia marina (L.) Besser Kure Atoll PRESERVED_SPECIMEN Lamoureux, C.H. 1910 Kure Atoll BISH 9/13/1961
203 Spergularia marina (L.) Besser honolulu, oahu. herb medicines, native name mimi ilio, (urine dog) PRESERVED_SPECIMEN pukui, m.k. Oahu BISH 8/8/1931
204 Spergularia marina (L.) Besser Summit trail south of Kealia, Oahu. Growing only on bare rocky clay of wide path PRESERVED_SPECIMEN Degener, O. 20670 Oahu BISH 3/14/1950
205 Spergularia marina (L.) Besser waimea, salt pond county park, w side of salt pan and adjacent edge of park Halephytic vegetation of Batis maritima and Pluchea indica PRESERVED_SPECIMEN lorence, d.h. collector number: 7438 Kauai BISH 2/5/1994
206 Spergularia marina (L.) Besser beach e of barber's point PRESERVED_SPECIMEN fosberg, f.r. collector number: 29528 Molokai BISH 2/4/1948
207 Spergularia marina (L.) Besser tern islet PRESERVED_SPECIMEN lamoureux, c.h. collector number: 1663 French Frigate Shoals BISH 9/2/1961
208 Spergularia marina (L.) Besser Eastern Island, edge of runway PRESERVED_SPECIMEN Herbst, D.R. 6409 Midway Atoll BISH 6/30/1980
209 Spergularia marina (L.) Besser Tern Islet, SE edge of runway PRESERVED_SPECIMEN Herbst, D.R. 6247 French Frigate Shoals BISH 9/11/1978
210 Spergularia marina (L.) Besser Summit trail S of Kealia bare rocky clay of wide path PRESERVED_SPECIMEN Degener, O. 20670 Oahu BISH 5/14/1950
211 Spergularia marina (L.) Besser summit trail south of kealia, oahu. growing only on bare rocky clay of wide path PRESERVED_SPECIMEN degener, o. collector number: 20670 Oahu BISH 3/14/1950
212 Spergularia marina (L.) Besser Beach near Koko Head PRESERVED_SPECIMEN Forbes, C.N. 1447 Oahu BISH 2/15/1909
213 Spergularia marina (L.) Besser Makai of aquaculture site of Kahuku Collected in solid, dwarfed strand community of Heliotropium anomalum, Jacquemontia, Fimbristylis, Nama, Euphorbia degeneri and Lycium. PRESERVED_SPECIMEN Stemmermann, R.L. 3703 Oahu BISH 2/11/1979
214 Spergularia marina (L.) Besser hamakua marsh, kailua, mauka (mountainside) of hamakua drive Low groundcover growing on dried mudflats with Bacopa, Paspalum. PRESERVED_SPECIMEN imada, c.t. engilis, a.; yuen, a.; dr.duk collector number: 92-6 Oahu BISH 4/4/1992
215 Spergularia marina (L.) Besser Sand Island, SE end of inner harbor mudflat PreservedSpecimen Herbst, D.R. Takeuchi, W.N. Collector Number: 6349 Midway Atoll BISH Specimen 6/29/1980
216 Spergularia marina (L.) Besser Eastern Island, edge of runway PreservedSpecimen Herbst, D.R. Collector Number: 6409 Midway Atoll BISH Specimen 6/30/1980
217 Spergularia marina (L.) Besser beach E of Barber's Point along shaded path in Prosopis on coral limestone PRESERVED_SPECIMEN Fosberg, F.R. 29528 Oahu BISH 2/4/1948
218 Spergularia marina (L.) Besser Ewa. Pohakupuna Rd PRESERVED_SPECIMEN Woolford, H.A. s.n. Oahu BISH
219 Spergularia marina (L.) Besser Near Koko Hd. on the beach PRESERVED_SPECIMEN Forbes, C.N. 1913.O Oahu BISH 7/1/1914
220 Spergularia marina (L.) Besser tern islet, se edge of runway PRESERVED_SPECIMEN herbst, d.r. collector number: 6247 French Frigate Shoals BISH 9/11/1978
221 Spergularia marina (L.) Besser Hamakua Marsh, Kailua, mauka (mountainside) of Hamakua Drive Low groundcover growing on dried mudflats with Bacopa, Paspalum. PRESERVED_SPECIMEN Imada, C.T. 92-6 Oahu BISH 4/4/1992
222 Spergularia marina (L.) Besser Waipio Peninsula; Waipio, Pouhala Marsh between Kapakahi & Waikele Streams Occasional herb on mudflats adjacent to Batis marshland & open water. PRESERVED_SPECIMEN Imada, C.T. 96-10 Oahu BISH 12/18/1996
223 Spergularia marina (L.) Besser s. of ewa, 1 mile w of oneula, in salt flats back of beach Weed PRESERVED_SPECIMEN st.john, h. collector number: 24804 Oahu BISH 4/27/1952
224 Spergularia marina (L.) Besser honolulu, north of ala moana park, in sand dredged from kewalo basin PRESERVED_SPECIMEN judd jr., a.f. Oahu BISH 5/25/1937
225 Spergularia marina (L.) Besser Kamilonui Valley, submerged in water by Kuapa Pond; bare soil; below Kiawe zone. (Near Koko Head.) PRESERVED_SPECIMEN Egler, F.E. 37-110 Oahu BISH 7/11/1937
226 Spergularia marina (L.) Besser near koko head. fish ponds outer brook PRESERVED_SPECIMEN bennett, t. collector number: 1501 Oahu BISH 6/11/1909
227 Spergularia marina (L.) Besser sand island, se end of inner harbor In dense rich forest. PRESERVED_SPECIMEN herbst, d.r. takeuchi, w.n. collector number: 6349 Hawaii BISH 6/29/1980
228 Spergularia marina (L.) Besser near haleiwa. marshy land PRESERVED_SPECIMEN hilliaurs Oahu BISH 3/17/1930
229 Spergularia marina (L.) Besser koko head oahu PRESERVED_SPECIMEN forbes, c.n. collector number: 1448 o. Oahu BISH 6/11/1909
230 Spergularia marina (L.) Besser Mokuleia Beach, Waialua Seasonally flooded bulldozed sand pit near ocean. PRESERVED_SPECIMEN Degener, O. 27846 Oahu BISH 2/1/1962
231 Spergularia marina (L.) Besser Pouhala Marsh, Waipahu Growing at edge and higher up on the bank of Pouhala. In wet to dry soil within the saltlines. Large patches in full sun to medium shade. PRESERVED_SPECIMEN Crago, L.M. Erickson, T. Collector Number: 2005-024 Oahu BISH 1/24/2005
232 Spergularia marina (L.) Besser e side of hale o lono harbor In sandy soil. PRESERVED_SPECIMEN oppenheimer, h.l. tangalin, n.; perlman, s.; garnett, b. collector number: h50629 Molokai BISH 5/15/2005
233 Spergularia marina (L.) Besser Near Haleiwa. Marshy land PRESERVED_SPECIMEN Hilliaurs s.n. Oahu BISH 3/17/1930
234 Spergularia marina (L.) Besser koko head. tuff sea cliffs opposite the koko head rifle range In small communities in exposed solution basins. Associated species: Heliotropium, Lipochaeta, Batis. PRESERVED_SPECIMEN nagata, k.m. collector number: 1232 Oahu BISH 7/25/1974
235 Spergularia marina (L.) Besser ewa, pohakupuna rd PRESERVED_SPECIMEN woolford, h.a. Oahu BISH 2/10/1957
236 Spergularia marina (L.) Besser near air field, puolo, hanapepe PRESERVED_SPECIMEN degener, o. degener, i. collector number: 27353 Kauai BISH 3/14/1961
237 Spergularia marina (L.) Besser barbers point, east of the lighthouse on dry thin soil back of the beach in the open PRESERVED_SPECIMEN egler, f.e. collector number: 37-274 Oahu BISH 8/11/1937
238 Spergularia marina (L.) Besser Near Koko Head. Fish Ponds outer brook PRESERVED_SPECIMEN Bennett, T. 1501 Oahu BISH 6/11/1909
239 Spergularia marina (L.) Besser Damp spot, Waipio PRESERVED_SPECIMEN Pearsall, G.A. s.n. Oahu BISH 3/19/1954
240 Spergularia marina (L.) Besser ewa, pohakupuna rd PRESERVED_SPECIMEN woolford, h.a. collector number: s.n. Oahu BISH 2/10/1957
241 Spergularia marina (L.) Besser near air field, Puolo, Hanapepe naturalized in mud flats PRESERVED_SPECIMEN Degener, O. 27353 Kauai BISH 3/14/1961
242 Spergularia marina (L.) Besser Ewa Plains Prosopis, Leucaena, Cynodon, Setaria, Many introduced plants. PRESERVED_SPECIMEN Higashino, P.K. 8189 Oahu BISH 2/11/1979
243 Spergularia marina (L.) Besser waipio peninsula; waipio, pouhala marsh between kapakahi & waikele streams Occasional herb on mudflats adjacent to Batis marshland & open water. PRESERVED_SPECIMEN imada, c.t. nishida, g. collector number: 96-10 Oahu BISH 12/18/1996
244 Spergularia marina (L.) Besser Koko Head. Tuff sea cliffs opposite the Koko Head Rifle Range In small but dense communities in solution basins. Associated with Batis, Heliotropium, Lipochaeta. PRESERVED_SPECIMEN Nagata, K.M. 1230 Oahu BISH 7/25/1974
245 Spergularia marina (L.) Besser Waimea, Salt Pond County Park, W side of salt pan and adjacent edge of park Halephytic vegetation of Batis maritima and Pluchea indica PRESERVED_SPECIMEN Lorence, D.H. 7438 Kauai BISH 2/5/1994
246 Spergularia marina (L.) Besser Sand Island, SE end of inner harbor mudflat PRESERVED_SPECIMEN Herbst, D.R. 6349 Midway Atoll BISH 6/29/1980
247 Spergularia marina (L.) Besser Sandspit in front of Kuapa Pond between Kuliouou and Portlock In sand just above high water mark PRESERVED_SPECIMEN Herbst, D.R. 413 Oahu BISH 4/5/1967
248 Spergularia marina (L.) Besser waialua, kawailua, roadside PRESERVED_SPECIMEN bush, w. topping, d.l. collector number: s.n. Oahu BISH 3/5/1933
249 Spergularia marina (L.) Besser Honolulu, Oahu. Herb medicines, native name MIMI ILIO, (urine dog) PRESERVED_SPECIMEN Pukui, M.K. s.n. Oahu BISH 4/14/1905
250 Spergularia marina (L.) Besser Ewa Beach PRESERVED_SPECIMEN Bennett, T. 174 Oahu BISH 2/24/1982
251 Spergularia marina (L.) Besser among branches of Atriplex sp. golden crown beard, mixed grasses, and Batis sp. west of Barbers Point Deep Draft harbor Plants were seeding and the population was robust. PRESERVED_SPECIMEN Funk, E.J. 403 Oahu BISH 5/1/1990
252 Spergularia marina (L.) Besser near koko head on the beach with j.f.s.stokes PRESERVED_SPECIMEN forbes, c.n. collector number: 1913.6 Oahu BISH 7/1/1914
253 Spergularia marina (L.) Besser pouhala marsh, waipahu Growing at edge and higher up on the bank of Pouhala. In wet to dry soil within the saltlines. Large patches in full sun to medium shade. PRESERVED_SPECIMEN crago, l.m. erickson, t. collector number: 2005-024 Oahu BISH 1/24/2005
254 Spergularia marina (L.) Besser Ca. 200 yds, east of the light house at Barber's Point. Arid, sandy region behind beach. Assoc. w/ Pluchea indica, P. odorata, Batis, Lycium, Sesuvium, Spergularia, sporobolus virg. Gregarious, naturalized. PRESERVED_SPECIMEN Herbst, D.R. 505 Oahu BISH 5/21/1967
255 Spergularia marina (L.) Besser kure atoll PRESERVED_SPECIMEN lamoureux, c.h. collector number: 1910 Kure Atoll BISH 9/13/1961
256 Spergularia marina (L.) Besser ewa. pohakupuna rd PRESERVED_SPECIMEN woolford, h.a. Oahu BISH
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